His M1 bike has been blighted by electronics issues all season.
Rossi had another race to forget in Misano and could only manage a seventh-place finish.
His team-mate, Maverick Vinales, performed moderately better and ended up in fifth spot.
The result was all the more embarrassing for Rossi in front of a passionate and partisan home crowd.
The frustrated veteran has urged Yamaha headquarters to get their house in order.
He said: “It was a pity because I would have liked to have a good race here in Misano, in my home Grand Prix, in front of all my friends and fans.
“I expected to be faster in the race, because yesterday in FP4 we did a good session.
“But this morning we had more problems with the bike, and even this afternoon, despite the conditions being more or less the same as those of yesterday.
“The big problem is that at the moment we don’t understand why.
“We must be able to understand what happens to our bike in situations like this.
“I hope we can fix it very quickly, because I want to go back to doing good races.”
The situation came to a head in qualifying before the Austrian Grand Prix last month.
Rossi started the race in 14th spot after a dismal Saturday session.
After the qualifying shambles, M1 project leader Kouji Tsuya publicly apologised to Rossi and Vinales.
BT Sport pundit James Toseland told Express Sport that huge oversights in Japan meant the heavyweights couldn’t bury their heads in the sand.
“I know Rossi has a lot of power in Yamaha,” Toseland exclusively told Express Sport.
“But for me there must have been a categorical mistake made in Japan that they’ve known about and it was too late to rectify.
“The results are showing the mistake, but obviously someone in Japan has made a wrong call.”
Tsuya is not a well known figure outside of Yamaha and Toseland says he was shocked to see the press conference called.
The former World Super Bike champion is also unconvinced it was the right call.
He added: “I think the public apology by Yamaha, that was such an unusual thing to see.
“There must have been some serious mistakes within the factory in Japan to allow one of their head developers to hold his hands up and apologise.
“I can’t remember any manufacturer ever doing that.
“I’m not sure if it was the right decision.
“If there were big mistakes made in Japan at Yamaha it’s very humble for them to hold their hands up and admit it in public.”
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